They use the same senses as we do and their feet and hands are similar to

 ours,　　　 chimpanzees still have opposable toes and can grasp things with their feet.

A.except what　　　　　　　　　　　　　B.except for

C.except that　　　　　　　　　　　　　D.besides that

They use the same senses as we do and their feet and hands are similar to ours, ________ chimpanzees still have opposable toes and can grasp things with their feet．

1. A.
   except what
2. B.
   except for
3. C.
   except that
4. D.
   besides that